Susan Middleton

Spineless: Portraits of Marine Invertebrates

Susan Middleton

Talk: Spineless: Portraits of Marine Invertebrates

Marine invertebrates make up more than 98% of the known animal species in the ocean, yet they remain elusive to most of us. Susan Middleton has devoted seven years of fieldwork across the Pacific Ocean to provide us with a rare glimpse into this mysterious world. As timely as it is visually stunning, her portraiture spotlights a vulnerable population, marine invertebrates being the first to experience the effects of climate change, particularly ocean acidification. From the stubby squid to the white phantom crab, these images open our eyes to the biodiversity, the fragility and the resiliency of these species. Photographs are copyright © Susan Middleton, all rights reserved

Susan Middleton

Susan Middleton is an acclaimed photographer, author, and lecturer specializing in the portraiture of rare and endangered animals, plants, sites, and cultures. For thirteen years she was the chair of the Department of Photography at the California Academy of Sciences, where she now serves as research associate.


In Spineless, acclaimed photographer Susan Middleton explores the mysterious and surprising world of marine invertebrates, which represent more than 98 percent of the known animal species in the ocean.

Find out more